My best guess is that, from the starting formation above, the caller wants the end of the center wave to work with one of the other center dancers (and, due to the call given, it would have to be the adjacent center dancer) plus the outside dancers that are in the clockwise direction.

Hence, from the starting formation above, I'd have couple #3 work with couple #4, and couple #1 work with couple #2.

This goes against my gestalt of jay, because I would want the two center dancers who are facing the same direction to work in the same jay.

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I have recently talked to the caller in question about this.

What the caller wanted (and should have said) was Clockwise "Z"s.

That is, each center pair of dancers work with the two outside positions that are clockwise to them, as the outside pairs work with the two inside positions that are clockwise to them. (similar to the Trace CONCEPT)